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Lawyer Garance Lombardi reunites with her ex’s family for a wedding. But when the bride is murdered, the past resurfaces. Garance quickly realizes that this murder signals the return of the Zodiac, who disappeared 20 years ago.

While Garance escapes from Eliott’s clutches, the revelation of Stella’s true identity sends shockwaves through the Escoffier family. As preparations for Juliette’s funeral are underway, Théo is kidnapped. Captain Keller, who tracked down the Zodiac twenty years ago, returns to duty and tries to unravel the Lombardi family’s secrets

Keller, Nadia, and Garance are doing everything they can to find Stella. Stella, who's Eliott’s captive, realizes that he is seeking revenge for his mother’s death. Keller, convinced that a mysterious mastermind is pulling the strings behind Eliott’s actions, begins to suspect Adrien. Nadia realizes that the Zodiac’s new crimes are linked to an old rape case.

The Lefur family is devastated by Fanny’s coma and Mathieu’s disappearance, as he is now the prime suspect. While Keller digs into the Escoffier family’s past, Nadia discovers that Mathieu has kidnapped Adrien. Garance realizes that she is the only one who can stop him from doing something irreversible

The Escoffier family is struck to the core. Secrets from the past resurface: Hélène confesses that she has done the unthinkable, while Keller and Nadia discover the full extent of Cyril Escoffier’s crimes. Garance decides to confront Audrey’s accusations, convinced that she is hiding something from her.

As ghosts from the past resurface and the Escoffier family faces yet another tragedy, Keller and Nadia discover that the source of the evil lies in Garance’s past. Stella confronts her past and uncovers her mother’s lies. The new Zodiac is about to carry out his revenge.