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Tony begins by asking why the village of St Bees has such a big church. Then in the Lake District, Tony discovers the origins of the scotch egg, before heading to England's most remote youth hostel.

Tony visits the Lake District, where he meets one of the last tenant sheep farmers, gets an exclusive tour of the only inhabited island on Derwentwater, learns how to cook chutney and gets a rare sighting of a red squirrel.

Tony visits Eden Valley, Cumbria ascending to the limestone-paved Great Asby Scar. He marvels at the engineering genius of the Smardale Gill viaduct, visits a dairy and investigates the prehistoric Nine Standards Rigg.

Tony explores the Yorkshire Dales, meeting the people who saved production of Wensleydale cheese in the town of Hawes, views the Aysgarth Falls and joins rehearsals with the Muker Silver Band.

In the North Yorkshire Moors Tony Robinson takes to the skies in a glider, admires the ruins of Rievaulx Abbey and learns about the heritage varieties of apples grown at Ampleforth Abbey.

In North Yorkshire, the presenter discovers how the longest stretch of steam railway would have disappeared, but for a team of volunteers dedicated to keeping the line and locomotives going. He also gets a rare glimpse inside the radar base at RAF Fylingdales, a missile-tracking facility that plays a significant role in the defence of both the UK and the USA. In Whitby, Tony visits the area's last shipyard and is delighted to hear that whales have returned to local waters after hunting almost drove them to extinction. His final destination is the picturesque coastal village of Robin Hood's Bay. Last in the series